Butterflies
and beetles

With around 2800 species in Upper Austria, butterflies are one of the most species-rich groups of animal organisms. The center of regional biodiversity can be found in the area of Kalkalpen National Park.
Butterfly with orange-brown colored wings sits on a white flower
Imperial Cloak ©ErichWEigand

Flying gemstones

Butterflies

Currently, 1560 butterfly species have been recorded and a further 200 species live in the immediately adjacent valleys of the national park. An internationally significant diversity, which is supported by partly unique and rare habitats! Due to the high degree of naturalness and the remarkable geomorphological and orographic conditions, there are important refugial areas for butterflies in the area, including a number of internationally protected or nationally endangered species.

Compared to other invertebrates, the level of knowledge about butterflies can already be regarded as relatively good. This above-average level of information is due to the traditional orientation of mainly voluntary, private researchers, some of whose research activities go back over 100 years. In particular, entomologists from the Steyr and Gmunden Entomological Working Groups have been very active since the establishment of the National Park: Herbert Kerschbaumsteiner (Styria), Dr. Martin Schwarz (Linz), Roland Mayrhofer (Steyr), August Pürstinger (Kirchdorf/Krems), Norbert Pöll (Bad Ischl), Franz Pühringer (St. Konrad) and unforgettably Josef Wimmer † (Steyr) and Friedrich Stöckl † (Vorchdorf) . Dr. Peter Huemer (Tyrolean Provincial Museum, Innsbruck) has published a popular scientific account of the special diversity of the local butterfly world in the book "Schmetterlinge - Vielfalt durch Wildnis".
In recent years, the butterfly fauna of the cultural landscape of the national park (alpine pastures, meadows) in particular has been studied and concrete management measures derived from this (Dr. Patrick Gros, Haus der Natur in Salzburg).

Two copper-colored butterflies copulate
Alpine mat fritillary butterfly © Erich Weigand
Cover page of book Butterflies Diversity through wildlife
Butterfly book © Kalkalpen National Park

Butterfly book for nature lovers

Butterflies - diversity through wildlife

Who would have thought that Kalkalpen National Park is home to more than 1,500 different butterfly species? This area of around 200 square kilometers is considered to be one of the last large wilderness areas in Central Europe, characterized by near-natural forests, unspoilt torrents and pristine mountain landscapes. Butterflies that have disappeared in many places find a last refuge here and many of them can still be observed frequently. From the gorge and alluvial forest near the valley, to alpine grass mats and rocky biotopes, to the gentle alpine pastures and meadows: Based on over 20 characteristic habitats, this book presents the typical butterfly fauna in each case. This makes the book a valuable companion for hikes in the Eastern Alps and especially in the Kalkalpen National Park area.

Authors: Peter Huemer, Peter Buchner, Josef Wimmer and Erich Weigand
Published by Verlag Trauner GmbH, 2014: ISBN978-3-99033-261-0

Alpine longhorned beetle flying off a piece of bark
Alpine longhorned beetle ©HerfriedMarek

Beetle

With around 7,400 species in Austria, beetles are the third most species-rich group of insects.

Many species are highly specialized and depend on very specific and small-scale habitats. A remarkably high number have specialized in forest habitats, particularly in the wood of dead trees. Such habitats can only be found in Central Europe in very remote areas that are difficult to manage in terms of forestry. However, it is particularly in these relict natural forest islands that most of the highly specialized beetle species, which experts refer to as "primeval forest relict species", have survived to this day. Many of these special rarities are native to Kalkalpen National Park and their population density is increasing due to the ever-growing forest wilderness with lots of dead wood. The best-known wood-dwelling beetle species in the national park is the alpine longhorn beetle. In addition to this, two other wood-dwelling nature conservation target species of the European Union with the highest protection category (Annex II of the Habitats Directive) can be found in the national park: the scarlet longhorned beetle and the mountain forest boring beetle.

Black-brown colored beetle with horn on its head crawls over a deadwood trunk
Horned skua ©ErichWeigand
Small brown beetle with long antennae sits on stone
Cave ground beetle © Erich Weigand

Arctaphaenops muellneri

Cave ground beetle

Müllner's Northeast Alpine blind beetle(Arctaphaenops muellneri)

The blind and almost transparent ground beetle is found exclusively in Upper Austria and is a true cave animal. It lives in perpetual darkness and is very fond of moisture. Almost the entire distribution area of this beetle lies within today's Kalkalpen National Park, which is why it also bears great responsibility for the conservation of this rarity. It is now known from five caves in Kalkalpen National Park and two more in the immediate vicinity.

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Small brown beetle with long antennae sits on stone
Cave ground beetle © Erich Weigand

Cave ground beetle Arctaphaenops muellneri

The cave ground beetle Arctaphaenops muellneri is one of the oldest and rarest animals observed in the Alps. It is one of the few surviving species that existed before several ice ages. Its known occurrence is limited to the Sengsengebirge and Reichraminger Hintergebirge. Species with such a limited occurrence are called endemics.

This beetle was discovered in 1970 in the Rettenbach Cave near Windischgarsten by the entomologist Karl Müllner from Steyr. The zoologist M.E. Schmid from Vienna was the first to document the special find as a species that was previously unknown worldwide and several million years old. Apart from the Rettenbach Cave, which was placed under protection as a natural monument due to this zoological peculiarity, only two other sites are known to date.

The six-millimetre cave ground beetle lives hidden in the widely branched, underground crevice system of the limestone mountains. It has adapted perfectly to these conditions. Its eyes and wings have regressed and its sun-protective pigments have also been lost due to its subterranean lifestyle. Instead, its sense of touch has developed considerably. Several long sensory bristles are distributed all over the body, the antennae are extremely long and have a joint at the base. Thorny appendages and long claws on the legs provide a secure grip in the dark. As a predator, the cave ground beetle is equipped with powerful mouthparts.

The evolutionary history of cave beetles is fascinating. In order to survive the warm periods, the beetles, which survived the Ice Age and are adapted to cool, damp conditions, had to find a suitable habitat niche. Wherever possible, such as on the edge of the north-eastern Limestone Alps, the animals found a refuge at higher altitudes. However, individual beetle populations became isolated as valleys were cut deeper and deeper. Over the course of thousands of years, several genetically independent species developed as a result, one of which is also found in today's Kalkalpen National Park.

Status, endangerment and protection

Status Kalkalpen NP: endemic, glacial relictStatus Austria (2005): endemic species
Red List Austria (2005): ----
Conservation responsibility Austria: particularly responsible
Need for action for Austria: not known
EU Habitats Directive: endemic status
Nature Conservation Act Upper Austria: protected

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