The Kalkalpen National Park
- an endemic kingdom

During the last ice age, the borders of today's Kalkalpen National Park lay on the edge of the great Alpine ice sheet. Many of the peaks in today's protected area remained at least partially ice-free. This enabled the well-adapted and above all cold-tolerant species to survive the cold periods in the area. The geographical isolation caused by the glaciation led to the emergence of new species over the course of thousands of years. As the occurrence of these species was often restricted to small areas, they are referred to as endemics (ancient Greek: "éndemos" native). Especially in the peripheral areas of the Alpine arc, such as the northern Limestone Alps, there is an increased occurrence of these unique ice age survivors. The national park and its surrounding mountain ranges are located right in the center of one of these "endemic hotspots".
Shiny metallic gold green ground beetle sits on dead wood
Golden shining ground beetle ©HerfriedMarek

Endemic beetle fauna

in the Kalkalpen National Park

With 174 known species and subspecies, the animal order of beetles (Coleoptera) is by far the largest group within Austria's endemic fauna. Of these, a very impressive 31 beetles (around 18 %) have also been recorded in Kalkalpen National Park , 13 of which are native only to Austria, without exception. For 18 others, at least 75 % of the distribution area is on Austrian territory. In addition, a further 14 endemic beetles have been confirmed for the protected area, at least 33% of whose total distribution area is on Austrian territory.

With a total of 45 known endemic species and a further 5 expected endemic species, Kalkalpen National Park and the Sengsengebirge in particular are among the most endemic areas in the Alps. Within Austria, due to their geographical location, only the higher mountain ranges in the immediate vicinity (especially Totes Gebirge, Haller Mauern, Gesäuse) have slightly higher numbers of species. Thus
the endemic beetle fauna alone shows that the Kalkalpen National Park fulfills its purpose as a protected area in the literal sense, i.e. as a refuge for sensitive and valuable species.

Probably the most important and also most famous endemic beetle of the Kalkalpen National Park is the cave ground beetle Arctaphaenops muellneri, whose distribution area (Sengsengebirge, Reichraminger Hintergebirge and Mollner Vorberge) lies primarily within the borders of the Kalkalpen National Park and thus exclusively in Upper Austria. A number of other locally endemic beetle species are only known from a few neighboring locations worldwide outside of the Kalkalpen National Park . These include Schauberger's weevil(Otiorhynchus schaubergeri), the extremely rare Austrian bearded beetle(Leistus austriacus) and the line-dotted beetle(Pterostichus lineatopunctatus).

The Kalkalpen National Park is a particularly important conservation area for these unique Austrian natural treasures. However, this also results in a high level of responsibility towards these specialists. The conservation of these species depends on the preservation of their habitats, but also on further research. Only if extensive knowledge about the special requirements, way of life and exact occurrence of a species is known can it be effectively protected and new challenges such as climate change or the accumulation of pollutants in the Alpine region be met.

Species list beetles endemics

List of 45 endemics detected in Kalkalpen National Park and 5 other endemics potentially expected for the protected area but not yet confirmed (1-48). Endemite criteria: Austrian endemic (E), subendemic (S), transnational endemic (T), and Alpine endemic (A). Conservation responsibility for Austria: (!) - to a high degree, (!!) - to a particularly high degree, (!!!) to an extremely high degree.

No.

Species/Subspecies

Family

German family

Status

Responsibility

1 Amara cuniculina Dejean, 1831 Carabidae Ground beetle E !!!
2 Arctaphaenops muellneri Schmid, 1972 Carabidae Ground beetle E !!!
3 Leistus austriacus Schauberger, 1925 Carabidae Ground beetle E !!!
4 Nebria hellwigii chalcicola Franz, 1949 Carabidae Ground beetle E !!!
5 Pterostichus lineatopunctatus L. Miller, 1850 Carabidae Ground beetle E !!!
6 Pterostichus selmanni selmanni (Duftschmid, 1812) Carabidae Ground beetle E !!!
7 Trechus ovatus ovatus Putzeys, 1846 Carabidae Ground beetle E !!!
8 Trechus wagneri Ganglbauer, 1906 Carabidae Ground beetle E !!!
9 Dichotrachelus vulpinus kraussi Penecke, 1894 Curculionidae Weevil E !!!
10 Otiorhynchus schaubergeri Lona, 1923 Curculionidae Weevil E !!!
11 Tropiphorus styriacus Bedel, 1883 Curculionidae Weevil E !!!
12 Aphodius montivagus Erichson, 1848 Scarabaeidae Leaf horned beetle E !!!
13 Leptusa gracilipes Krauss, 1899 Staphylinidae Short-winged beetle E !!!
14 Carabus alpestris alpestris Sturm, 1815 Carabidae Ground beetle S !!
15 Carabus auronitens intercostatus Gredler, 1854 Carabidae Ground beetle S !!
16 Carabus sylvestris haberfelneri Ganglbauer, 1891 Carabidae Ground beetle S !!
17 Nebria germari norica Schauberger, 1927 Carabidae Ground beetle S !!
18 Oreonebria austriaca (Ganglbauer, 1889) Carabidae Ground beetle S !!
19 Pterostichus illigeri (Panzer, 1803) Carabidae Ground beetle S !!
20 Pterostichus panzeri (Panzer, 1803) Carabidae Ground beetle S !!
21 Pterostichus subsinuatus (Dejean, 1828) Carabidae Ground beetle S !!
22 Trechus alpicola alpicola Sturm, 1825 Carabidae Ground beetle S !!
23 Trechus hampei Ganglbauer, 1891 Carabidae Ground beetle S !!
24 Trechus pinkeri Ganglbauer, 1891 Carabidae Ground beetle S !!
25 Trechus rotundipennis (Duftschmid, 1812) Carabidae Ground beetle S !!
26 Oreina plagiata commutata (Suffrian, 1861) Chrysomelidae Leaf beetle S !!
27 Psylliodes subaeneus styriacus Heikertinger, 1921 Chrysomelidae Leaf beetle S !!
28 Brachiodontus alpinus (Hampe, 1867) Curculionidae Weevil S !!
29 Otiorhynchus nocturnus Reitter, 1913 Curculionidae Weevil S !!
30 Otiorhynchus pigrans Stierlin, 1861 Curculionidae Weevil S !!
31 Rhinomias austriacus (Reitter, 1894) Curculionidae Weevil S !!
32 Hydraena alpicola Pretner, 1931 Hydraenidae Long-tailed water beetle S !!
33 Lathrobium testaceum Kraatz, 1857 Staphylinidae Short-winged beetle S !!
34 Malthodes subductus Kiesenwetter, 1863 Cantharidae Soft-bodied beetle T !
35 Carabus fabricii fabricii Duftschmid, 1812 Carabidae Ground beetle T !
36 Patrobus styriacus Chaudoir, 1871 Carabidae Ground beetle T !
37 Pterostichus jurinei jurinei (Panzer, 1803) Carabidae Ground beetle T !
38 Pterostichus transversalis (Duftschmid, 1812) Carabidae Ground beetle T !
39 Euconnus carinthiacus Ganglbauer, 1896 Scydmaenidae Short-winged beetle T !
40 Aleochara ganglbaueri Bernhauer, 1901 Staphylinidae Short-winged beetle T !
41 Bryaxis bothrophorus (Stolz, 1917) Staphylinidae Short-winged beetle T !
42 Oreonebria castanea (Bonelli, 1810) Carabidae Ground beetle A !
43 Aleochara meschniggi Bernhauer, 1901 Staphylinidae Short-winged beetle A !
44 Eusphalerum nitidicolle (Baudi, 1857) Staphylinidae Short-winged beetle A !
45 Leptusa globulicollis (Mulsant & Rey, 1853) Staphylinidae Short-winged beetle A !
46 Leptusa piceata (Mulsant & Rey, 1853) Staphylinidae Short-winged beetle A !
47 Ocypus brevipennis (Heer, 1839) Staphylinidae Short-winged beetle A !

Confirmed primeval forest relic beetle species

Species list of the beetle fauna

Data: A. Eckelt, H. Mitter & E. Weigand (2023); number of records: Locations in Kalkalpen National Park, without indication of the number of individuals observed

No.

primeval forest-Relict types (scientific)

Family (scientific)

Family (German)

Cat.

First detection NP

Evidence

1 Ampedus carpathicus (=suecicus) Elateridae Click beetle 1 2019 2
2 Bius thoracicus Tenebrionidae Black beetle 1 2004 3
3 Bolitophagus interruptus Tenebrionidae Black beetle 1 2019 1
4 Calitys scabra Trogossitidae Hunting beetle, flat beetle 1 1982 6
5 Ischnodes sanguinicollis Elateridae Click beetle 1 2019 1
6 Lacon lepidopterus Elateridae Click beetle 1 2018 3
7 Peltis grossa Trogossitidae Hunting beetle, flat beetle 1 2010 27
8 Phryganophilus ruficollis Melandryidae Darkling beetle 1 2019 3
9 Stictoleptura erythroptera Cerambycidae Longhorn beetle 1 2019 1
10 Allecula rhenana Tenebrionidae Black beetle 2 2019 2
11 Ampedus auripes Elateridae Click beetle 2 2010 3
12 Ampedus melanurus Elateridae Click beetle 2 2019 2
13 Cerophytum elateroides Cerophytidae click beetle-like 2 2019 1
14 Ceruchus chrysomelinus Lucanidae Stag beetle 2 1977 12
15 Crepidophorus mutilatus Elateridae Click beetle 2 2015 1
16 Cryptophagus lysholmi Cryptophagidae Mold beetle 2 2011 1
17 Derodontus macularis Derodontidae Larch beetle 2 2015 1
18 Dicerca berolinensis Buprestidae jewel beetle 2 2019 7
19 Dircaea australis Melandryidae Darkling beetle 2 2011 7
20 Ernobius explanatus Anobiidae Poching beetle 2 2010 3
21 Gnorimus variabilis Scarabaeidae Leaf horned beetle 2 2019 1
22 Grynocharis oblonga Trogossitidae Hunting beetle, flat beetle 2 2019 1
23 Hexarthrum duplicatum Curculionidae Weevil 2 2015 1
24 Latridius brevicollis Latridiidae Fashion beetle 2 2010 1
25 Leiestes seminiger Endomychidae Squirrel beetle 2 2019 1
26 Lordithon speciosus Staphylinidae Short winged 2 2010 2
27 Mycetochara obscura Tenebrionidae Black beetle 2 2018 1
28 Mycetoma suturale Tetratomidae Cull beetle 2 2015 2
29 Mycetophagus decempunctatus Mycetophagidae Dry rot beetle 2 2019 1
30 Nematodes filum Eucnemidae Comb beetle 2 2009 7
31 Phymatura brevicollis Staphylinidae Short winged 2 2012 1
32 Prionychus melanarius Tenebrionidae Black beetle 2 2012 5
33 Prostomis mandibularis Prostomidae Shovel beetle 2 2011 8
34 Quedius truncicola Staphylinidae Short winged 2 2019 2
35 Rhizophagus brancsiki Monotomidae Root beetle 2 2019 1
36 Rosalia alpina Cerambycidae Longhorn beetle 2 1968 25
37 Stephanopachys substriatus Bostrichidae Boring beetle 2 2004 1
38 Synchita separanda Zopheridae Bark beetle 2 2010 7
39 Tragosoma depsarium Cerambycidae Longhorn beetle 2 1985 3
40 Triplax collaris Erotylidae Fungus beetle 2 2019 1
41 Xestobium austriacum Anobiidae Poching beetle 2 2010 6
An orange-brown colored butterfly with wings sitting open on a pink flower
Ash fritillary butterfly on flower ©ErichWeigand

Endangered butterfly species

Endangered species in Austria with occurrence in Kalkalpen National Park

Listing of selected species occurring in Kalkalpen National Park which are listed as endangered (VU), critically endangered (EN) or even threatened with extinction (CR) in the Red List of endangered butterflies and moths in Austria (excluding moths). The number of previous records in the national park area is also listed.

German species name

Red List Austria

Evidence

Large Papperglucke CR 1
Yellow ringed butterfly EN 91
Ash fritillary EN 49
Light gray gray owl EN 6
Heilziest butterfly EN 5
Bedstraw hawkmoth EN 1
Small hop rootworm EN 1
Gray fashionable wood owl EN 1
Scalloped bear VU 48
Augsburg bear VU 43
Elegant daisy VU 31
Plum jelly VU 30
Purple bear VU 14
Platinum gray tufted owl VU 5
Sunflower fritillary butterfly VU 5
Rockland burrowing owl VU 5
Elm tip butterfly VU 4
Plague root rot VU 3
Spotted fritillary VU 3
Meadow dusty owl VU 2
Ruderal Field Scops Owl VU 2
Large kingfisher VU 2
Simulans ground owl VU 2
Alexis blue, sky-blue Seinklee blue VU 1
Gray lichen owl VU 1
Dry grassland ground owl VU 1
Common knapweed VU 1
Small sainfoin blue VU 1
Round-spotted cube fritillary VU 1
Rocky scrubland burrowing owl VU 1
Sainfoin Oriole VU 1
Plantain Fritillary VU 1
Small spring snail with red-brown shell
Macro photo spring snail ©ErichWeigand

Endemic spring snail species

The primary aim of this project was to identify the species of spring snails (hydrobioids: dwarf snails) found in the area, the majority of which are endemic. The majority of this species group, which is widespread in karst spring biotopes and underground waters in the Northern Limestone Alps, cannot be reliably identified to species level on the basis of morphological characteristics and requires genetic analyses. However, knowledge of the species status is essential, as endemic species are usually distributed on an extremely small scale and the majority are also under strict conservation protection. The genetic analysis as well as other classical methods for species determination were carried out under a contract for work by employees of the Natural History Museum Vienna (laboratory & ABOL project management: Dr. Luise Kruckenhauser, head of the mollusc collection and project coordination: Anita Eschner, project implementation: Hannah Schubert, as well as the involved experts Dr. Michael Duda and Dr. Martin Haase).

100 DNA barcodes were created on individuals from 39 spring biotopes. The results confirm that extremely rare hydrobioids, which are particularly worthy of protection due to their endemic status, occur in selected spring biotopes in the region of the national park and therefore require special protection. This applies in particular to the two species Belgrandiella aulaei and Bythiospeum nocki, which have so far only been recorded worldwide in Kalkalpen National Park and its immediate surroundings.

Spring snail species (scientific)

 
Belgrandiella aulaei  
Bythiospeum nocki  
Bythinella concia  
Hauffenia kerschneri  
Hauffenia wienerwaldensis  
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